Carole A. Cheah

Carole Cheah

Valley Laboratory
The Connecticut Agricultural Experiment Station
153 Cook Hill Road
Windsor, CT 06095

Voice: (860) 683-4980 Fax: (860) 683-4987
E-mail: Carole.Cheah@ct.gov


Expertise:
I am trained in the research, application, assessment and implementation of the theory and practice of biological control of invasive agricultural and forest pests and weeds. My areas of expertise center on the evaluation of attributes of natural enemies as potential candidates for biological control of the invasive insect pests, hemlock woolly adelgid and elongate hemlock scale threatening the sustainability of eastern hemlock, Tsuga canadensis. My expertise is in the detailed study of the biology, behavior and field adaptation of biological control agents such as the Japanese coccinellid, Sasajiscymnus (=Pseudoscymnus) tsugae, and other predator species, and on the development of improved laboratory mass-rearing methods for them. I also conduct surveys and long-term field assessments of forest hemlock health to monitor the effects of biological control in the context of changing climatic and environmental conditions. I am the author and recipient of over 12 major federal grants from the USDA Forest Service and USDA APHIS for such research and employment over the past decade as a durational research entomologist with The CAES.

Education:
Ph.D. Wolfson College, University of Cambridge, England 1988
M. Phil. (Applied Entomology), Wolfson College, University of Cambridge 1984
B.A. (Hons.) Zoology, University of Oxford, England 1982

Station Career:
Assistant Agricultural Scientist II 2003- present
Assistant Agricultural Scientist I 2000-2003
Postdoctoral Research Scientist 1994-2000

Past Research:
I have worked on numerous projects evaluating the potential of biological control agents, parasitoids and predators, for pests such as the chrysanthemum leaf-miner, hemlock woolly adelgid (HWA), elongate hemlock scale (EHS) and mile-a-minute weed (MAM).  Many of the predator species originate from Asia and I have experience working in the federal quarantine laboratory of the USDA Forest Service.  Extensive collaborations with Dr. Allen Cohen of North Carolina State University have resulted in the development of a novel and superior artificial dietary supplement for enhancing the survival and longevity of diverse predators imported for biological control of HWA and MAM.  Together with Dr. Cohen, we have also developed a series of process and production quality control tests for improving the laboratory rearing of predators for biological control of HWA.

Current Research:
I am continuing research into the evaluation of plant volatiles as potential attractants for improved field monitoring of Sasajiscymnus tsugae, a Japanese ladybeetle predator of HWA discovered by CAES; development of laboratory methods for culturing native predators of EHS for augmentative biological control, and continuation of the implementation and monitoring in Connecticut of the imported weevil, Rhinoncomimus latipes, for biological control of MAM.

Selected publications available from the author, Carole.Cheah@ct.gov:

Cheah, C. (2021).Collaborative biological control in Connecticut with Sasajiscymnus tsugae, introduced predator of Hemlock Woolly Adelgid (HWA). Connecticut Agricultural Experiment Station Bulletin 1081. https://portal.ct.gov/-/media/CAES/DOCUMENTS/Publications/Bulletins/B1081.pdf

 

Cheah, C. (2019).Hemlock Woolly Adelgid webpage for the ConnecticutAgricultural Experiment Station at https://portal.ct.gov/CAES/Publications/Publications/Hemlock-Woolly-Adelgid

Cheah, C. A. S-J. and Ellis, D. R. (2019). Spread of Mile-a-minute vine, Persicaria perfoliata L. (Polgonaceae) to Connecticut islands in Long Island Sound.

Rhodora, 121(987), 219-221. DOI: 10.3119/19-02

 

Cheah, C. (2019). Climate Impacts on Eastern Hemlock Sustainability. The Habitat, 31(1), 6-7, 13-14. PDF: https://portal.ct.gov/CAES/Publications/Publications/Hemlock-Woolly-Adelgid

Cheah, C. (2019).Hemlock woolly adelgid (HWA) and other factors impacting eastern hemlock. Connecticut Agricultural Experiment Station Fact Sheet.

PDF: https://portal.ct.gov/-/media/CAES/DOCUMENTS/Publications/Fact_Sheets/Valley_Laboratory/Hemlock-Woolly-Adelgid-Factsheet-Dec-2018-Cheah-Final.pdf?la=en

Cheah, C. (2019). Battling Mile-a-minute Weed in Connecticut. The Habitat, 31(2), 6-7.

Cheah, C.A.S.-J. (2017).Predicting winter mortality of hemlock woolly adelgid in Connecticut by climatic divisions. Northeastern Naturalist, 24(Special Issue 7), DOI: 10.1656/045.024.s713 PDF: https://portal.ct.gov/CAES/Publications/Publications/Hemlock-Woolly-Adelgid

 

Cheah, C. (2016).  HWA winter mortality in Connecticut: implications for management and control. Connecticut Agricultural Experiment Station Fact Sheet. PDF:

http://www.ct.gov/caes/lib/caes/documents/publications/fact_sheets/plant_pathology and_ecology/hemlock_woolly_adelgid_winter_mortality__7.12.16.pdf

Cheah, C. and Li, D. (2016). The Red Bark Phenomenon. Connecticut Agricultural

Experiment Station Fact Sheet. Available online at:

http://www.ct.gov/caes/lib/caes/documents/publications/fact_sheets/valley_laboratory/the_red_bark_phenomenon.pdf

Russo, N. J., Cheah, C.A. S-J., and Tingley, M. W. (2016). Experimental evidence for branch-to-bird transfer as a mechanism of avian dispersal of the hemlock woolly adelgid (Hempitera: Adelgidae). Environmental Entomology, 45, 1107-1114. https://doi.org/10.1093/ee/nvw083

Cohen, A. C and Cheah, C. A.S.-J. (2015). Interim Diets for Specialist Predators of Hemlock Woolly Adelgids. Entomol. Ornithol. Herpetol. 4, 153. DOI:10.4172/2161-0983.1000153

https://www.omicsonline.org/open-access/interim-diets-for-specialist-predators-of-hemlock-woolly-adelgids-2161-0983-1000153.php?aid=52757

Cohen, A. C. and Cheah, C. (2011).Chap. 14. Development of artificial diets for predators of hemlock woolly adelgids. In: Implementation and Status of Biological Control of Hemlock Woolly Adelgid. Tech. Coor. Onken, B. and Reardon, R. USDA Forest Service FHTET Publication FHTET-2011-04 pp. 148-160.

Cohen, A. C., Cheah, C., Kidd, K. and Hodgson, T. (2011). Chap. 13. Defining PC/QC standards of mass-rearing HWA predators. In: Implementation and Status of Biological Control of Hemlock Woolly Adelgid. Tech. Coor. Onken, B. and Reardon, R. USDA Forest Service FHTET Publication FHTET-2011-04 pp. 139-147.

Cheah, C. (2011).  Chapter 4. Sasajiscymnus (=Pseudoscymnustsugae, a ladybeetle from Japan. In: Implementation and Status of Biological Control of Hemlock Woolly Adelgid. Tech.Coor. Onken, B. and Reardon, R. USDA Forest Service FHTET Publication FHTET-2011-04 pp. 43-52.

Cohen, A. C. and Cheah, C. (2010). Packaging and Presentation of Artificial Diets for Hemlock Woolly Adelgid Predators In: Proceedings of the Fifth HWA Symposium in the Eastern United States, Asheville, NC, August 17-19. Compilers Onken, B. and  Reardon, R.  USDA Forest Service FHTET-2010-07. p 33-35.

Cheah, C. (2010).Connecticut’s threatened landscape: natural enemies of biological control of invasive species. Frontiers of Plant Science57(2), 5-16

Cheah, C.A.S-J. and McClure, M. S. (2010). Sasajiscymnus (formerly Pseudoscymnustsugae (Coleoptera: Coccinellidae). In: Biological Control: A Guide to Natural Enemies in North America. A. Shelton. Cornell University College of Agriculture and Life Sciences, Department of Entomology website. http://www.nysaes.cornell.edu/ent/biocontrol/predators/sasajiscymnus.html

Cohen, A. C., Cheah, C. A. S-J., Strider, J., and Hain, F. (2008)Diet development for hemlock woolly adelgids and their predators In: Proceedings of the Fourth Symposium on the Hemlock Woolly Adelgid in the Eastern United States, Feb.12-14, 2008, Hartford, Connecticut. Onken, B. and Reardon, R. (Compilers) FHTET 2008-01 p.150-156.

Cheah, C. (2008). The case for Sasajiscymnus tsugae: Biological control has helped save Connecticut’s hemlocks.  Abstract for a poster in Proceedings of the Fourth Symposium on Hemlock Woolly Adelgid in the Eastern United States. Hartford, CT.  Feb. 12-14, 2008. Onken, B. O. and Reardon, R. (Compilers) FHTET 2008-01. pp. 279-280. PDF: https://portal.ct.gov/CAES/Publications/Publications/Hemlock-Woolly-Adelgid

Cohen, A. C., Cheah, C. A. S.-J., Strider, J., and Hain, F. (2008). Diet development for hemlock woolly adelgids and their predators. In: Proceedings of the Fourth Symposium on the Hemlock Woolly Adelgid in the Eastern United States, February 12-14, 2008, Hartford, Connecticut. Onken, B. and Reardon, R.  (Compilers) FHTET 2008-01 p. 150-156. PDF: https://portal.ct.gov/CAES/Publications/Publications/Hemlock-Woolly-Adelgid

Cheah, C. (2006). Hope for hemlocks. Connecticut Woodlands, 71(3), 13-15. PDF: https://portal.ct.gov/CAES/Publications/Publications/Hemlock-Woolly-Adelgid

Cheah, C. A. S-J., Mayer, M. A., Palmer, D., Scudder, T. and Chianese, R. (2005).  Assessments of biological control of hemlock woolly adelgid with Sasajiscymnus tsugae in Connecticut and New Jersey.  In: Proceedings of the Third Symposium on the Hemlock Woolly Adelgid in the Eastern United States, Feb.1-3, 2005, Asheville, North Carolina. Onken, B. and Reardon, R. (Compilers) FHTET 2005-01 pp. 116-130.

Cheah, C, Montogmery, M. Salom, S. Parker, B. L., Costa, S. and Skinner, M. (2004). Biological control of hemlock woolly adelgid. USDA Forest Service. FHTET-2002-04, Reardon, R. and B. Onken (Tech. Coordinators), 22 pp.

Cheah, C. A. S-J. and McClure, M. S. (2002).Pseudoscymnus tsugae in Connecticut forests: the first five years. Proceedings of the Hemlock Woolly Adelgid Symposium, East Brunswick, NJ, February 5-7.  Eds. Onken, B., Reardon, R. Lashomb, J.  p. 150-165.

McClure, M. S. and Cheah, C. A. S-J. (2002). Important mortality factors in the life cycle of hemlock woolly adelgid, Adelges tsugae Annand (Homptera:  Adelgidae) in the Northeastern United States. Onken, B., Reardon, and R. Lashomb, J. (Eds.). Proceedings of the Hemlock Woolly Adelgid Symposium, East Brunswick, NJ, February 5-7. p. 13-22.

Cheah, C. A.S-J. and McClure, M. S.(2000). Seasonal synchrony between the exotic predator, Pseudoscymnus tsugae (Coleoptera: Coccinellidae) and its prey, the hemlock woolly adelgid, Adelges tsugaeAgriculture and Forest Entomology, 2, 241-251

McClure, M. S. and Cheah, C. A. S-J. (1999). Reshaping the ecology of invading populations of hemlock woolly adelgid, Adelges tsugae, (Homoptera:  Adelgidae) in eastern North America. Biological Invasions, 1, 247-254.

Cheah, C. A. S-J. and McClure, M. S. (1998). Life history and development of Pseudoscymnus tsugae (Coleoptera: Coccinellidae), a new predator of the hemlock woolly adelgid (Homoptera: Adelgidae).  Environmental Entomology27, 1531-1536.

Coaker, T. H. and Cheah, C. A. (1993). Conditioning as a factor in parasitoid host plant preference. Biocontrol Science and Technology, 3, 277-283.

Cheah, C. A. and Coaker, T. H. (1992). Host finding and discrimination in Diglyphus isaea, a parasitoid of the chrysanthemum leaf miner, Chromatomyia syngenesiae. Biocontrol Science and Technology, 2, 109-118.

Cheah, C. S-J. (1987). Temperature requirements of the chrysanthemum leaf miner, Chromatomyia syngenesiae (Dipt.: Agromyzidae), and its ectoparasitoid, Digylyphus isaea (Hym.:Eulophiodae). Entomophaga32, 357-365